Nurhachius luei, a new istiodactylid pterosaur (Pterosauria, Pterodactyloidea) from the Early Cretaceous Jiufotang Formation of Chaoyang City, Liaoning Province (China) and comments on the Istiodactylidae [PeerJ]
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Last updated 11 abril 2025
![Nurhachius luei, a new istiodactylid pterosaur (Pterosauria, Pterodactyloidea) from the Early Cretaceous Jiufotang Formation of Chaoyang City, Liaoning Province (China) and comments on the Istiodactylidae [PeerJ]](https://dfzljdn9uc3pi.cloudfront.net/2019/7688/1/fig-1-full.png)
A new istiodactylid pterosaur, Nurhachius luei sp. nov., is here reported based on a complete skull with mandible and some cervical vertebrae from the lower part of the Jiufotang Formation of western Liaoning (China). This is the second species of Nurhachius, the type-species being N. ignaciobritoi from the upper part of the Jiufotang Formation. A revised diagnosis of the genus Nurhachius is provided, being this taxon characterized by the presence of a slight dorsal deflection of the palatal anterior tip, which is homoplastic with the Anhangueria and Cimoliopterus. N. luei sp. nov. shows an unusual pattern of tooth replacement, with respect to other pterodactyloid species. The relationships within the Istiodactylidae and with their closest taxa are investigated through a phylogenetic analysis by parsimony.
![Nurhachius luei, a new istiodactylid pterosaur (Pterosauria, Pterodactyloidea) from the Early Cretaceous Jiufotang Formation of Chaoyang City, Liaoning Province (China) and comments on the Istiodactylidae [PeerJ]](https://www.tandfonline.com/cms/asset/b505034b-5714-4f81-b7fc-b8cadf2ba887/ghbi_a_2222127_f0003_oc.jpg)
Full article: A new pterosaur specimen from the Lower Cretaceous Yixian Formation of Liaoning Province, China: The oldest fossil record of Nurhachius
![Nurhachius luei, a new istiodactylid pterosaur (Pterosauria, Pterodactyloidea) from the Early Cretaceous Jiufotang Formation of Chaoyang City, Liaoning Province (China) and comments on the Istiodactylidae [PeerJ]](https://upload.wikimedia.org/wikipedia/commons/f/f9/Caulkicephalus_trimicrodon.jpg)
Caulkicephalus - Wikipedia
![Nurhachius luei, a new istiodactylid pterosaur (Pterosauria, Pterodactyloidea) from the Early Cretaceous Jiufotang Formation of Chaoyang City, Liaoning Province (China) and comments on the Istiodactylidae [PeerJ]](https://d3i71xaburhd42.cloudfront.net/487b13f43836ff28f088688fe1644a8425a38c4b/9-Figure4-1.png)
PDF] An unusual long-tailed pterosaur with elongated neck from western Liaoning of China.
![Nurhachius luei, a new istiodactylid pterosaur (Pterosauria, Pterodactyloidea) from the Early Cretaceous Jiufotang Formation of Chaoyang City, Liaoning Province (China) and comments on the Istiodactylidae [PeerJ]](https://palaeo-electronica.org/content/images/1015/s_figure3.jpg)
A new istiodactylid pterosaur
![Nurhachius luei, a new istiodactylid pterosaur (Pterosauria, Pterodactyloidea) from the Early Cretaceous Jiufotang Formation of Chaoyang City, Liaoning Province (China) and comments on the Istiodactylidae [PeerJ]](https://dfzljdn9uc3pi.cloudfront.net/2023/14829/1/fig-8-2x.jpg)
A taxonomic revision of the Sinopterus complex (Pterosauria, Tapejaridae) from the Early Cretaceous Jehol Biota, with the new genus Huaxiadraco [PeerJ ]
![Nurhachius luei, a new istiodactylid pterosaur (Pterosauria, Pterodactyloidea) from the Early Cretaceous Jiufotang Formation of Chaoyang City, Liaoning Province (China) and comments on the Istiodactylidae [PeerJ]](https://d3i71xaburhd42.cloudfront.net/62dab115295dec24dc704597fbe9e790cb624dc5/12-Figure5-1.png)
PDF] Taxonomy of the Lonchodectidae (Pterosauria, Pterodactyloidea)
![Nurhachius luei, a new istiodactylid pterosaur (Pterosauria, Pterodactyloidea) from the Early Cretaceous Jiufotang Formation of Chaoyang City, Liaoning Province (China) and comments on the Istiodactylidae [PeerJ]](https://d3i71xaburhd42.cloudfront.net/001199e0972964c9af58f79ae72d0003c5ed8364/5-Figure1-1.png)
PDF] A new specimen of Sinopterus dongi (Pterosauria, Tapejaridae) from the Jiufotang Formation (Early Cretaceous, China)
![Nurhachius luei, a new istiodactylid pterosaur (Pterosauria, Pterodactyloidea) from the Early Cretaceous Jiufotang Formation of Chaoyang City, Liaoning Province (China) and comments on the Istiodactylidae [PeerJ]](https://upload.wikimedia.org/wikipedia/commons/a/ae/Pterodactylus_holotype_fly_mmartyniuk.png)
Timeline of pterosaur research - Wikipedia
![Nurhachius luei, a new istiodactylid pterosaur (Pterosauria, Pterodactyloidea) from the Early Cretaceous Jiufotang Formation of Chaoyang City, Liaoning Province (China) and comments on the Istiodactylidae [PeerJ]](https://reptileevolution.com/images/lepidosauromorpha/diadectidae/lepidosauriformes/fenestrasauria/pterosauria/nurhachius2_recon588.gif)
Nurhachius
![Nurhachius luei, a new istiodactylid pterosaur (Pterosauria, Pterodactyloidea) from the Early Cretaceous Jiufotang Formation of Chaoyang City, Liaoning Province (China) and comments on the Istiodactylidae [PeerJ]](https://d3i71xaburhd42.cloudfront.net/001199e0972964c9af58f79ae72d0003c5ed8364/11-Figure5-1.png)
PDF] A new specimen of Sinopterus dongi (Pterosauria, Tapejaridae) from the Jiufotang Formation (Early Cretaceous, China)
![Nurhachius luei, a new istiodactylid pterosaur (Pterosauria, Pterodactyloidea) from the Early Cretaceous Jiufotang Formation of Chaoyang City, Liaoning Province (China) and comments on the Istiodactylidae [PeerJ]](https://upload.wikimedia.org/wikipedia/commons/thumb/3/3c/Nurhachius_sp._skull_NMNS.jpg/220px-Nurhachius_sp._skull_NMNS.jpg)
Nurhachius - Wikipedia
![Nurhachius luei, a new istiodactylid pterosaur (Pterosauria, Pterodactyloidea) from the Early Cretaceous Jiufotang Formation of Chaoyang City, Liaoning Province (China) and comments on the Istiodactylidae [PeerJ]](https://dfzljdn9uc3pi.cloudfront.net/2019/7688/1/fig-5-full.png)
Nurhachius luei, a new istiodactylid pterosaur (Pterosauria, Pterodactyloidea) from the Early Cretaceous Jiufotang Formation of Chaoyang City, Liaoning Province (China) and comments on the Istiodactylidae [PeerJ]
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